![Picture of Alun](https://www.mmu.ac.uk/sites/default/files/styles/page_header_half/public/2022-08/AW__3.jpg?h=1bcd3856&itok=eZCcY892)
Professor Alun Williams PhD FBASES FTPS
Professor of Sport and Exercise Genomics
I conduct research into and teach the genetics of physical performance, injury risk, including how that relates to sex categories in sport.
My profile
Biography
I’m Professor of Sport and Exercise Genomics and my main research interest is the genetics of human physical performance and health. I hold honorary research positions at University College London and Swansea University, and I’ve been elected as a Fellow of the British Association of Sport and Exercise Sciences and as a Fellow of The Physiological Society.
My current research projects investigate the genetic characteristics of elite sport performance, and the physical changes that occur during testosterone suppression in trans women. I regularly publish and review for leading peer-reviewed journals in my field and also advise sports governing bodies regarding sex category eligibility policies.
I’ve written several position stands / expert statements on scientific and ethical issues in sport, covering scientific and ethical issues related to the use of genetic technology in sport and exercise, as well as the complex and controversial issue of eligibility for sex categories in sport. I also regularly speak at conferences and other events on these topics.
Interests and expertise
My area of expertise lies broadly in the upper limits of human physical performance. I have published widely on genetic profiles of elite athletes, the improvements in performance that can result from physical training, and sport-related injuries, as well as ethical and policy issues. Some of my current areas of interest focus on elite athletes in several sports, the genetic basis of concussion injury, and the role of testosterone on physical performance in different groups such as athletes with differences in sex development (DSDs) and transgender athletes.
Impact
I have served as independent expert witness at the Court of Arbitration for Sport. I’ve written several consensus statements, and advise sports governing bodies, regarding sex and age categories in sport, and the use of genetic technology in sport. I communicate widely regarding the often misleading messages communicated by genetic testing companies about their products available to the general public and elite sports organisations.
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Genetics and ethics collide at the pinnacle of sport
Read moreTeaching
I’ve supervised many research students to successful completions in sport science, genetics and exercise physiology. I’m always willing to consider applications for PhDs and other higher degrees from promising students.
I also teach on our MSc and BSc programmes, focussing on genetics in sport and exercise, as well as other related aspects of exercise physiology.
Research outputs
Please see the lists of outputs below. Most of these should be accessible via various platforms, but drop me a line if you would like to access anything.
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Chapters in books
Massidda, M., Calo, C.M., Cieszczyk, P., Kikuchi, N., Ahmetov, I.I., Williams, A.G. (2019) 'Genetics of team sports.' In Barh, D., Ahmetov, I.I. (ed.) Sports, Exercise and Nutritional Genomics. London, UK: Elsevier, pp. 105-128.
Moran, C.N., Williams, A.G., Wang, G. (2019) 'Using elite athletes as a model for genetic research.' Routledge Handbook of Sport and Exercise Systems Genetics. pp. 357-371.
Wang, G., Tanaka, M., Eynon, N., North, K.N., Williams, A.G., Collins, M., Moran, C.N., Britton, S.L., Fuku, N., Ashley, E.A., Klissouras, V., Lucia, A., Ahmetov, I.I., De Geus, E., Alsayrafi, M., Pitsiladis, Y.P. (2016) 'The Future of Genomic Research in Athletic Performance and Adaptation to Training.' pp. 55-67.
Williams, A.G., Wackerhage, H., Day, S.H. (2016) 'Genetic Testing for Sports Performance, Responses to Training and Injury Risk: Practical and Ethical Considerations.' pp. 105-119.
Bradburn, S., McPhee, J.S., Williams, A.G., Heffernan, S.M., Lockey, S.J., Day, S.H., Murgatroyd, C. (2015) 'Protocol for multiplex amplicon sequencing using barcoded primers.' In Slevin, M., McDowell, G. (ed.) Handbook of Vascular Biology Techniques. Springer, pp. 427-438.
Day, S.H., Williams, A.G. (2012) 'Genetics, trainability.' Day, S.H. & Williams, A.G. Genetics, Trainability. In F.C. Mooren, ed. Encyclopedia of Exercise Medicine in Health and Disease. Berlin: Springer, 2012. Springer,
Williams, A.G., Wackerhage, H. (2009) 'Genetic testing of athletes..' pp. 176-186.
Little, T., Williams, A.G. (2005) 'Specificity of acceleration, maximum speed and agility in professional soccer players.' In Reilly, T., Cabri, J., Aroujo, D. (ed.) Science and Football V: The Proceedings of the Fifth World Congress on Science and Football. Oxon, UK: Routledge, pp. 276-283.
Little, T., Williams, A.G. (2005) 'The effects of different modes of stretching during warm-up on high speed motor capacities in professional footballers.' In Reilly, T., Cabri, J., Araujo, D. (ed.) Science and Football V: The Proceedings of the Fifth World Congress on Science and Football. Oxon, UK: Routledge, pp. 380-386.
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Reports
Webborn, N., Williams, A.G., McNamee, M., Bouchard, C., Pitsiladis, Y., Ahmetov, I., Ashley, E., Byrne, N., Camporesi, S., Collins, M., Dijkstra, P., Eynon, N., Fuku, N., Garton, F., Hoppe, N., Holm, S., Kaye, J., Klissouras, V., Lucia, A., Maase, K., Moran, C., North, K.N., Pigozzi, F., Wang, G. (2015) Consensus Statement on direct-to-consumer genetic testing for sports performance and talent identification.
Williams, A.G., Miah, A., Harris, R.C., Montgomery, H.E., Wackerhage, H. (2012) The BASES Expert Statement on genetic research and testing in sport and exercise science.
Williams, A.G., Wackerhage, H., Miah, A., Harris, R.C., Montgomery, H.E. (2007) Genetic research and testing in sport and exercise science: BASES Position Stand. Leeds, UK:
Williams, A.G., Rayson, M.P. (1997) Improved Physical Training Enhances Military Performance in Recruits: Report for the Ministry of Defence.
Williams, A.G., Rayson, M.P. (1997) The Optimisation of Training during CMS(R): Report for the Ministry of Defence.
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Journal articles
Lundberg, T.R., Tucker, R., McGawley, K., Williams, A.G., Millet, G.P., Sandbakk, Ø., Howatson, G., Brown, G.A., Carlson, L.A., Chantler, S., Chen, M.A., Heffernan, S.M., Heron, N., Kirk, C., Murphy, M.H., Pollock, N., Pringle, J., Richardson, A., Santos-Concejero, J., Stebbings, G.K., Christiansen, A.V., Phillips, S.M., Devine, C., Jones, C., Pike, J., Hilton, E.N. (2024) 'The International Olympic Committee framework on fairness, inclusion and nondiscrimination on the basis of gender identity and sex variations does not protect fairness for female athletes.' Scandinavian Journal of Medicine and Science in Sports, 34(3)
Shaw, A.L., Williams, A.G., Stebbings, G.K., Chollier, M., Harvey, A., Heffernan, S.M. (2024) 'The perspective of current and retired world class, elite and national athletes on the inclusion and eligibility of transgender athletes in elite sport.' Journal of Sports Sciences, 42(5) pp. 381-391.
Fraser, K.K., Williams, A.G., de Silva, T.T.A., Stebbings, G.K., Backhouse, S.H. (2023) 'Making competent decisions in sport and exercise science and sports medicine: preliminary practical guidelines on sex and gender.' Journal of Science and Medicine in Sport, 27(4) pp. 281-284.
Murtagh, C.F., Hall, E.C., Brownlee, T.E., Drust, B., Williams, A.G., Erskine, R.M. (2023) 'The genetic association with athlete status, physical performance and injury risk in soccer.' International Journal of Sports Medicine, 44(13) pp. 941-960.
Antrobus, M.R., Brazier, J., Callus, P., Herbert, A.J., Stebbings, G.K., Day, S.H., Kilduff, L.P., Bennett, M.A., Erskine, R.M., Raleigh, S.M., Collins, M., Pitsiladis, Y.P., Heffernan, S.M., Williams, A.G. (2023) 'Concussion-associated gene variant COMT rs4680 is associated with elite rugby athlete status.' Clinical Journal of Sport Medicine, 33(5)
Brazier, J., Antrobus, M.R., Herbert, A.J., Callus, P.C., Khanal, P., Stebbings, G.K., Day, S.H., Heffernan, S.M., Kilduff, L.P., Bennett, M.A., Erskine, R.M., Raleigh, S.M., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2023) 'Gene Variants Previously Associated with Reduced Soft-tissue Injury Risk: Part 2 - polygenic associations with elite status in rugby.' European Journal of Sport Science, 23(8) pp. 1779-1788.
Hall, E.C.R., Lockey, S.J., Heffernan, S.M., Herbert, A.J., Stebbings, G.K., Day, S.H., Collins, M., Pitsiladis, Y.P., Erskine, R.M., Williams, A.G. (2023) 'The PPARGC1A Gly482Ser polymorphism is associated with elite long-distance running performance.' Journal of Sports Sciences, 41(1) pp. 56-62.
Dines, H.R., Nixon, J., Lockey, S.J., Herbert, A.J., Kipps, C., Pedlar, C.R., Day, S.H., Heffernan, S.M., Antrobus, M.R., Brazier, J., Erskine, R.M., Stebbings, G.K., Hall, E.C.R., Williams, A.G. (2022) 'Collagen Gene Polymorphisms Previously Associated with Resistance to Soft-Tissue Injury Are More Common in Competitive Runners Than Nonathletes.' Journal of Strength and Conditioning Research, 37(4) pp. 799-805.
Khanal, P., Morse, C.I., He, L., Herbert, A.J., Onambélé-Pearson, G.L., Degens, H., Thomis, M., Williams, A.G., Stebbings, G.K. (2022) 'Polygenic Models Partially Predict Muscle Size and Strength but Not Low Muscle Mass in Older Women.' Genes, 13(6) pp. 982-982.
Antrobus, M.R., Brazier, J., Callus, P.C., Herbert, A.J., Stebbings, G.K., Khanal, P., Day, S.H., Kilduff, L.P., Bennett, M.A., Erskine, R.M., Raleigh, S.M., Collins, M., Pitsiladis, Y.P., Heffernan, S.M., Williams, A.G. (2022) 'Concussion-Associated Polygenic Profiles of Elite Male Rugby Athletes.' Genes, 13(5) pp. 820-820.
Hall, E.C.R., Larruskain, J., Gil, S.M., Lekue, J.A., Baumert, P., Rienzi, E., Moreno, S., Tannure, M., Murtagh, C.F., Ade, J.D., Squires, P., Orme, P., Anderson, L., Whitworth-Turner, C.M., Morton, J.P., Drust, B., Williams, A.G., Erskine, R.M. (2022) 'Injury risk is greater in physically mature versus biologically younger male soccer players from academies in different countries.' Physical Therapy in Sport, 55(May) pp. 111-118.
Brazier, J., Antrobus, M.R., Herbert, A.J., Callus, P.C., Stebbings, G.K., Day, S.H., Heffernan, S.M., Kilduff, L.P., Bennett, M.A., Erskine, R.M., Raleigh, S.M., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2022) 'Gene Variants Previously Associated with Reduced Soft Tissue Injury Risk: Part 1 - Independent Associations with Elite Status in Rugby.' European Journal of Sport Science, 23(5) pp. 726-735.
Hall, E.C.R., Larruskain, J., Gil, S.M., Lekue, J.A., Baumert, P., Rienzi, E., Moreno, S., Tannure, M., Murtagh, C.F., Ade, J.D., Squires, P., Orme, P., Anderson, L., Whitworth-Turner, C.M., Morton, J.P., Drust, B., William, A.G., Erskine, R.M. (2022) 'Playing Position is Associated with Injury Incidence Rate in Male Academy Soccer Players.' Journal of Athletic Training, 57(7) pp. 696-703.
Kikuchi, N., Moreland, E., Homma, H., Semenova, E.A., Saito, M., Larin, A.K., Kobatake, N., Yusupov, R.A., Okamoto, T., Nakazato, K., Williams, A.G., Generozov, E.V., Ahmetov, I.I. (2021) 'Genes and Weightlifting Performance.' Genes, 13(1) pp. 25-25.
Khanal, P., Williams, A.G., He, L., Stebbings, G.K., Onambele-Pearson, G.L., Thomis, M., Degens, H., Morse, C.I. (2021) 'Sarcopenia, obesity, and sarcopenic obesity: Relationship with skeletal muscle phenotypes and single nucleotide polymorphisms.' Journal of Clinical Medicine, 10(21)
Herbert, A.J., Williams, A.G., Lockey, S.J., Erskine, R.M., Sale, C., Hennis, P.J., Day, S.H., Stebbings, G.K. (2021) 'Correction to: Bone mineral density in high-level endurance runners: part A—site-specific characteristics.' European Journal of Applied Physiology, 122(1) pp. 269-271.
Hall, E.C., Baumert, P., Larruskain, J., Gil, S.M., Lekue, J.A., Rienzi, E., Moreno, S., Tannure, M., Murtagh, C.F., Ade, J.D., Squires, P., Orme, P., Anderson, L., Brownlee, T.E., Whitworth-Turner, C.M., Morton, J.P., Drust, B., Williams, A.G., Erskine, R.M. (2021) 'The genetic association with injury risk in male academy soccer players depends on maturity status.' Scandinavian journal of medicine & science in sports, 32(2) pp. 338-350.
Herbert, A.J., Williams, A.G., Lockey, S.J., Erskine, R.M., Sale, C., Hennis, P.J., Day, S.H., Stebbings, G.K. (2021) 'Bone mineral density in high-level endurance runners: Part B—genotype-dependent characteristics.' European Journal of Applied Physiology, 122(1) pp. 71-80.
Herbert, A.J., Williams, A.G., Lockey, S.J., Erskine, R.M., Sale, C., Hennis, P.J., Day, S.H., Stebbings, G.K. (2021) 'Bone mineral density in high-level endurance runners: part A—site-specific characteristics.' European Journal of Applied Physiology, 121(12) pp. 3437-3445.
Khanal, P., He, L., Degens, H., Stebbings, G.K., Onambele-Pearson, G.L., Williams, A.G., Thomis, M., Morse, C.I. (2021) 'Dietary protein requirement threshold and micronutrients profile in healthy older women based on relative skeletal muscle mass.' Nutrients, 13(9)
Hall, E.C.R., Almeida, S.S., Heffernan, S.M., Lockey, S.J., Herbert, A.J., Callus, P., Day, S.H., Pedlar, C.R., Kipps, C., Collins, M., Pitsiladis, Y.P., Bennett, M.A., Kilduff, L.P., Stebbings, G.K., Erskine, R.M., Williams, A.G. (2021) 'Genetic Polymorphisms Related to VO2max Adaptation Are Associated With Elite Rugby Union Status and Competitive Marathon Performance.' International Journal of Sports Physiology and Performance, 16(12) pp. 1858-1864.
Khanal, P., He, L., Stebbings, G.K., Onambele-Pearson, G.L., Degens, H., Williams, A.G., Thomis, M., Morse, C.I. (2021) 'Static one-leg standing balance test as a screening tool for low muscle mass in healthy elderly women.' Aging Clinical and Experimental Research, 33(7) pp. 1831-1839.
Antrobus, M.R., Brazier, J., Stebbings, G.K., Day, S.H., Heffernan, S.M., Kilduff, L.P., Erskine, R.M., Williams, A.G. (2021) 'Genetic factors that could affect concussion risk in elite rugby.' Sports, 9(2) pp. 19-19.
Varley, I., Stebbings, G., Williams, A.G., Day, S., Hennis, P., Scott, R., Grazette, N., Herbert, A.J. (2021) 'An investigation into the association of bone characteristics and body composition with stress fracture in athletes.' The Journal of Sports Medicine and Physical Fitness, 61(11) pp. 1490-1498.
Khanal, P., He, L., Herbert, A.J., Stebbings, G.K., Onambele-Pearson, G.L., Degens, H., Morse, C.I., Thomis, M., Williams, A.G. (2020) 'The Association of Multiple Gene Variants with Ageing Skeletal Muscle Phenotypes in Elderly Women.' Genes, 11(12) pp. 1459-1459.
Moreland, E., Borisov, O.V., Semenova, E.A., Larin, A.K., Andryushchenko, O.N., Andryushchenko, L.B., Generozov, E.V., Williams, A.G., Ahmetov, I.I. (2020) 'Polygenic Profile of Elite Strength Athletes.' Journal of Strength and Conditioning Research, 36(9) pp. 2509-2514.
He, L., Khanal, P., Morse, C.I., Williams, A., Thomis, M. (2020) 'Associations of combined genetic and epigenetic scores with muscle size and muscle strength: a pilot study in older women.' Journal of Cachexia, Sarcopenia and Muscle, 11(6) pp. 1548-1561.
Hall, E., Murgatroyd, C., Stebbings, G., Cunniffe, B., Harle, L., Salter, M., Ramadass, A., Westra, J., Hunter, E., Akoulitchev, A., Williams, A. (2020) 'The Prospective Study of Epigenetic Regulatory Profiles in Sport and Exercise Monitored Through Chromosome Conformation Signatures.' Genes, 11(8)
Hall, E.C.R., Larruskain, J., Gil, S.M., Lekue, J.A., Baumert, P., Rienzi, E., Moreno, S., Tannure, M., Murtagh, C.F., Ade, J.D., Squires, P., Orme, P., Anderson, L., Whitworth-Turner, C.M., Morton, J.P., Drust, B., Williams, A.G., Erskine, R.M. (2020) 'An injury audit in high-level male youth soccer players from English, Spanish, Uruguayan and Brazilian academies.' Physical Therapy in Sport, 44pp. 53-60.
Khanal, P., He, L., Stebbings, G., Onambele-Pearson, G.L., Degens, H., Williams, A., Thomis, M., Morse, C.I. (2020) 'Prevalence and association of single nucleotide polymorphisms with sarcopenia in older women depends on definition.' Scientific Reports, 10(1) pp. 2913-2913.
He, L., Khanal, P., Morse, C., Williams, A., Thomis, M. (2019) 'Differentially methylated gene patterns between age-matched sarcopenic and non-sarcopenic women.' Journal of Cachexia, Sarcopenia and Muscle, 10(6) pp. 1295-1306.
Brazier, J., Antrobus, M., Stebbings, G., Day, S., Heffernan, S., Cross, M., Williams, A. (2019) 'Tendon and ligament injuries in elite rugby: The potential genetic influence.' Sports, 7(6) pp. 138-165.
Herbert, A.J., Williams, A.G., Hennis, P.J., Erskine, R.M., Sale, C., Day, S.H., Stebbings, G.K. (2019) 'The interactions of physical activity, exercise and genetics and their associations with bone mineral density: implications for injury risk in elite athletes.' European Journal of Applied Physiology, 119(1) pp. 29-47.
Brazier, J., Antrobus, M., Stebbings, G., Day, S., Callus, P., Erskine, R., Bennett, M.A., Kilduff, L.P., Williams, A.G. (2018) 'The anthropometric and physiological characteristics of elite rugby athletes.' Journal of Strength and Conditioning Research, 34(6) pp. 1790-1781.
Stebbings, G.K., Williams, A.G., Herbert, A.J., Lockey, S.J., Heffernan, S.M., Erskine, R.M., Morse, C.I., Day, S.H. (2018) 'TTN genotype is associated with fascicle length and marathon running performance..' Scandinavian Journal of Medicine and Science in Sports, 28(2)
Papadimitriou, I.D., Lockey, S.J., Voisin, S., Herbert, A.J., Garton, F., Houweling, P.J., Cieszczyk, P., Maciejewska-Skrendo, A., Sawczuk, M., Massidda, M., Calò, C.M., Astratenkova, I.V., Kouvatsi, A., Druzhevskaya, A.M., Jacques, M., Ahmetov, I.I., Stebbings, G.K., Heffernan, S., Day, S.H., Erskine, R., Pedlar, C., Kipps, C., North, K.N., Williams, A.G., Eynon, N. (2018) 'No association between ACTN3 R577X and ACE I/D polymorphisms and endurance running times in 698 Caucasian athletes..' BMC Genomics, 19(1)
Varley, I., Patel, S., Williams, A.G., Hennis, P.J. (2018) 'The current use, and opinions of elite athletes and support staff in relation to genetic testing in elite sport within the UK.' Biology of Sport, 35(1) pp. 13-19.
Heffernan, S.M., Kilduff, L.P., Erskine, R.M., Day, S.H., Stebbings, G.K., Cook, C.J., Raleigh, S.M., Bennett, M.A., Wang, G., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2017) 'COL5A1 gene variants previously associated with reduced soft tissue injury risk are associated with elite athlete status in rugby.' BMC Genomics, 18(Suppl 8)
Stebbings, G.K., Day, S.H., Williams, A., Morse, C. (2017) 'Polymorphisms in PTK2 are associated with skeletal muscle specific force: an independent replication study.' European Journal of Applied Physiology, 117(4) pp. 713-720.
Heffernan, S.M., Stebbings, G.K., Kilduff, L.P., Erskine, R.M., Day, S.H., Morse, C.I., McPhee, J.S., Cook, C.J., Vance, B., Ribbans, W.J., Raleigh, S.M., Roberts, C., Bennett, M.A., Wang, G., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2017) 'Fat mass and obesity associated (FTO) gene influences skeletal muscle phenotypes in non-resistance trained males and elite rugby playing position..' BMC Genetics, 18(1) pp. 4-4.
Hussain, A.W., Onambele, G.L., Williams, A.G., Morse, C.I. (2016) 'Medial gastrocnemius specific force of adult men with spastic cerebral palsy.' Muscle & Nerve, 56(2) pp. 298-306.
Boyd, C., Eaves, S.J., Morse, C., Roach, N., Williams, A.G., Barnes, C. (2016) 'A time-motion analysis of paralympic football for athletes with cerebral palsy.' International Journal of Sports Science & Coaching, 11(4) pp. 552-558.
Pitsiladis, Y.P., Tanaka, M., Eynon, N., Bouchard, C., North, K.N., Williams, A.G., Collins, M., Moran, C.N., Britton, S.L., Fuku, N., Ashley, E.A., Klissouras, V., Lucia, A., Ahmetov, I.I., de Geus, E., Alsayrafi, M. (2016) 'Athlome Project Consortium: a concerted effort to discover genomic and other "omic" markers of athletic performance..' Physiological Genomics, 48
Heffernan, S.M., Kilduff, L.P., Erskine, R.M., Day, S.H., McPhee, J.S., McMahon, G.E., Stebbings, G.K., Neale, J.P., Lockey, S.J., Ribbans, W.J., Cook, C.J., Vance, B., Raleigh, S.M., Roberts, C., Bennett, M.A., Wang, G., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2016) 'Association of ACTN3 R577X but not ACE I/D gene variants with elite rugby union player status and playing position..' Physiological Genomics, 48
Webborn, N., Williams, A.G., McNamee, M., Bouchard, C., Pitsiladis, Y., Ahmetov, I., Ashley, E., Byrne, N., Camporesi, S., Collins, M., Dijkstra, P., Eynon, N., Fuku, N., Garton, F.C., Hoppe, N., Holm, S., Kaye, J., Klissouras, V., Lucia, A., Maase, K., Moran, C., North, K.N., Pigozzi, F., Wang, G. (2015) 'Direct-to-consumer genetic testing for predicting sports performance and talent identification: Consensus statement.' British Journal of Sports Medicine, 49
Heffernan, S.M., Kilduff, L.P., Day, S.H., Pitsiladis, Y.P., Williams, A.G. (2015) 'Genomics in rugby union: A review and future prospects.' European Journal Of Sport Science, 15
Foster, B.P., Morse, C.I., Onambele, G.L., Williams, A.G. (2015) 'Erratum to: Variants within the MMP3 gene and patellar tendon properties in vivo in an asymptomatic population (vol 114, pg 2625, 2014).' European Journal of Applied Physiology, 115(4) pp. 861-861.
Foster, B.P., Morse, C.I., Onambele, G.L., Williams, A.G. (2014) 'Variants within the MMP3 gene and patellar tendon properties in vivo in an asymptomatic population.' European Journal of Applied Physiology, 114
Erskine, R.M., Williams, A.G., Jones, D.A., Stewart, C.E., Degens, H. (2014) 'The individual and combined influence of ACE and ACTN3 genotypes on muscle phenotypes before and after strength training..' Scand J Med Sci Sports, 24(4) pp. 642-648.
Ateş, F., Yucesoy, C.A. (2014) 'Effects of botulinum toxin type A on non‐injected bi‐articular muscle include a narrower length range of force exertion and increased passive force.' Muscle & Nerve, 49(6) pp. 866-878.
Erskine, R.M., Morse, C.I., Day, S.H., Williams, A.G., Onambele-Pearson, G.L. (2014) 'The human patellar tendon moment arm assessed in vivo using dual-energy X-ray absorptiometry.' Journal of biomechanics, 47(6) pp. 1294-1298.
Foster, B.P., Morse, C.I., Onambele, G.L., Williams, A.G. (2014) 'Human COL5A1 rs12722 gene polymorphism and tendon properties in vivo in an asymptomatic population.' European Journal of Applied Physiology, 114(7) pp. 1393-1402.
Williams, A.G., Heffernan, S.M., Day, S.H. (2014) 'Genetic testing in exercise and sport - have direct-to-consumer genetic tests come of age?.' Science and Sport: Current Trends, 2(1) pp. 3-9.
Hussain, A.W., Onambele, G.L., Williams, A.G., Morse, C.I. (2014) 'Muscle size, activation, and coactivation in adults with cerebral palsy..' Muscle Nerve, 49(1) pp. 76-83.
Williams, A.G., Day, S.H., Lockey, S.J., Heffernan, S.M., Erskine, R.M. (2014) 'Genomics as a practical tool in sport - have we reached the starting line?.' Cellular and Molecular Exercise Physiology, 3
Stebbings, G.K., Morse, C.I., Williams, A.G., Day, S.H. (2014) 'Variability and distribution of muscle strength and its determinants in humans..' Muscle Nerve, 49(6) pp. 879-886.
Hussain, A.W., Onambele, G.L., Williams, A.G., Morse, C.I. (2013) 'Passive stiffness of the gastrocnemius muscle in athletes with spastic hemiplegic cerebral palsy..' Eur J Appl Physiol, 113(9) pp. 2291-2299.
Gabbasov, R.T., Arkhipova, A.A., Borisova, A.V., Hakimullina, A.M., Kuznetsova, A.V., Williams, A.G., Day, S.H., Ahmetov, I.I. (2013) 'The HIF1A gene Pro582Ser polymorphism in Russian strength athletes..' J Strength Cond Res, 27(8) pp. 2055-2058.
Eynon, N., Nasibulina, E.S., Banting, L.K., Cieszczyk, P., Maciejewska-Karlowska, A., Sawczuk, M., Bondareva, E.A., Shagimardanova, R.R., Raz, M., Sharon, Y., Williams, A.G., Ahmetov, I.I., Lucia, A., Birk, R. (2013) 'The FTO A/T Polymorphism and Elite Athletic Performance: A Study Involving Three Groups of European Athletes.' PLoS ONE, 8(4)
Wang, G., Padmanabhan, S., Wolfarth, B., Fuku, N., Lucia, A., Ahmetov, I.I., Cieszczyk, P., Collins, M., Eynon, N., Klissouras, V., Williams, A., Pitsiladis, Y. (2013) 'Genomics of elite sporting performance: What little we know and necessary advances.' 84pp. 123-149.
Erskine, R.M., Williams, A.G., Jones, D.A., Stewart, C.E., Degens, H. (2012) 'Do PTK2 gene polymorphisms contribute to the interindividual variability in muscle strength and the response to resistance training? A preliminary report.' JOURNAL OF APPLIED PHYSIOLOGY, 112(8) pp. 1329-1334.
Dhamrait, S.S., Williams, A.G., Day, S.H., Skipworth, J., Payne, J.R., World, M., Humphries, S.E., Montgomery, H.E. (2012) 'Variation in the uncoupling protein 2 and 3 genes and human performance.' Journal of Applied Physiology, 112(7) pp. 1122-1127.
Foster, B.P., Morse, C.I., Onambele-Pearson, G., Ahmetov, I.I., Williams, A.G. (2012) 'Genetic variation, protein composition and potential influences on tendon properties in humans.' The open sports medicine journal, 6(1) pp. 8-21.
Williams, A.G., Miah, A., Harris, R.C., Montgomery, H.E., Wackerhage, H. (2012) 'The BASES Expert Statement on genetic research and testing in sport and exercise science.' The Sport and Exercise Scientist, 31(Spring) pp. 14-15.
Ahmetov, I.I., Vinogradova, O.L., Williams, A.G. (2012) 'Gene polymorphisms and fiber-type composition of human skeletal muscle.' International Journal of Sport Nutrition and Exercise Metabolism, 22(4) pp. 292-303.
Hughes, D.C., Day, S.H., Ahmetov, I.I., Williams, A.G. (2011) 'Genetics of muscle strength and power: polygenic profile similarity limits skeletal muscle performance..' J Sports Sci, 29(13) pp. 1425-1434.
McPhee, J.S., Perez-Schindler, J., Degens, H., Tomlinson, D., Hennis, P., Baar, K., Williams, A.G. (2011) 'HIF1A P582S gene association with endurance training responses in young women..' Eur J Appl Physiol, 111(9) pp. 2339-2347.
McPhee, J.S., Williams, A.G., Perez-Schindler, J., Degens, H., Baar, K., Jones, D.A. (2011) 'Variability in the magnitude of response of metabolic enzymes reveals patterns of co-ordinated expression following endurance training in women..' Exp Physiol, 96(7) pp. 699-707.
Stellingwerff, T., Jeukendrup, A.E. (2011) 'Don't forget the gut--it is an important athletic organ!.' Journal of applied physiology (Bethesda, Md. : 1985), 110(1) pp. 278-294.
Ahmetov, I.I., Druzhevskaya, A.M., Lyubaeva, E.V., Popov, D.V., Vinogradova, O.L., Williams, A.G. (2011) 'The dependence of preferred competitive racing distance on muscle fibre type composition andACTN3genotype in speed skaters.' Experimental Physiology, 96(12) pp. 1302-1310.
Williams, A.G. (2011) 'The two-hour marathon: Genetics will become increasingly important.' Journal of Applied Physiology, 110(1)
Erskine, R.M., Jones, D.A., Maffulli, N., Williams, A.G., Stewart, C.E., Degens, H. (2011) 'What causes in vivo muscle specific tension to increase following resistance training?.' Exp Physiol, 96(2) pp. 145-155.
Williams, A.G. (2011) 'Athleticogenomics, epigenetics and gene doping.' Highlights, 19(2) pp. 12-15.
Erskine, R.M., Jones, D.A., Williams, A.G., Stewart, C.E., Degens, H. (2010) 'Inter-individual variability in the adaptation of human muscle specific tension to progressive resistance training..' Eur J Appl Physiol, 110(6) pp. 1117-1125.
McPhee, J.S., Williams, A.G., Degens, H., Jones, D.A. (2010) 'Inter-individual variability in adaptation of the leg muscles following a standardised endurance training programme in young women..' Eur J Appl Physiol, 109(6) pp. 1111-1118.
Erskine, R.M., Jones, D.A., Williams, A.G., Stewart, C.E., Degens, H. (2010) 'Resistance training increases in vivo quadriceps femoris muscle specific tension in young men..' Acta Physiol (Oxf), 199(1) pp. 83-89.
Dinsdale, N.J., Williams, A.G. (2010) 'Can forefoot varus wedges enhance anaerobic cycling performance in untrained males with forefoot varus?.' Sport Scientific and Practical Aspects, 7(2) pp. 5-10.
Wackerhage, H., Miah, A., Harris, R.C., Montgomery, H.E., Williams, A.G. (2009) 'Genetic research and testing in sport and exercise science: A review of the issues.' Journal of Sports Sciences, 27(11) pp. 1109-1116.
Ahmetov, I.I., Williams, A.G., Popov, D.V., Lyubaeva, E.V., Hakimullina, A.M., Fedotovskaya, O.N., Mozhayskaya, I.A., Vinogradova, O.L., Astratenkova, I.V., Montgomery, H.E., Rogozkin, V.A. (2009) 'The combined impact of metabolic gene polymorphisms on elite endurance athlete status and related phenotypes.' Human Genetics, 126(6) pp. 751-761.
Ahmetov, I.I., Popov, D.V., Lyubaeva, E.V., Missina, S.S., Vinogradova, O.L., Hakimullina, A.M., Rogozkin, V.A., Williams, A.G. (2009) 'Association of the VEGFR2 gene His472Gln polymorphism with endurance-related phenotypes.' European Journal of Applied Physiology, 107(1) pp. 95-103.
McPhee, J.S., Williams, A.G., Stewart, C., Baar, K., Schindler, J.P., Aldred, S., Maffulli, N., Sargeant, A.J., Jones, D.A. (2009) 'The training stimulus experienced by the leg muscles during cycling in humans..' Exp Physiol, 94(6) pp. 684-694.
Williams, A.G., Wackerhage, H. (2009) 'Genetic research and testing in sport and exercise science: A summary of the BASES position stand.' Deutsche Zeitschrift fur Sportmedizin, 60(10) pp. 331-335.
Williams, A.G., Folland, J.P. (2008) 'Reply from Alun G. Williams and Jonathan P. Folland.' Journal of Physiology, 586(12) pp. 3019-3020.
Folland, J.P., Mc Cauley, T.M., Williams, A.G. (2008) 'Allometric scaling of strength measurements to body size.' European Journal of Applied Physiology, 102(6) pp. 739-745.
Williams, A.G., Folland, J.P. (2008) 'Similarity of polygenic profiles limits the potential for elite human physical performance..' J Physiol, 586(1) pp. 113-121.
Folland, J.P., Williams, A.G. (2007) 'Methodological issues with the interpolated twitch technique.' Journal of Electromyography and Kinesiology, 17(3) pp. 317-327.
LITTLE, T., WILLIAMS, A.G. (2007) 'EFFECTS OF SPRINT DURATION AND EXERCISE.' Journal of Strength and Conditioning Research, 21(2) pp. 646-648.
Little, T., Williams, A.G. (2007) 'Effects of sprint duration and exercise: Rest ratio on repeated sprint performance and physiological responses in professional soccer players.' Journal of Strength and Conditioning Research, 21(2) pp. 646-648.
Williams, A.G., Wilkinson, M. (2007) 'Simple anthropometric and physical performance tests to predict maximal box-lifting ability..' J Strength Cond Res, 21(2) pp. 638-642.
Little, T., Williams, A.G. (2007) 'Measures of exercise intensity during soccer training drills with professional soccer players.' Journal of Strength and Conditioning Research, 21(2) pp. 367-371.
Folland, J.P., Williams, A.G. (2007) 'Calculation of muscle activation using neuromuscular electrical stimulation [3].' Medicine and Science in Sports and Exercise, 39(4)
Folland, J.P., Williams, A.G. (2007) 'The adaptations to strength training: Morphological and neurological contributions to increased strength.' Sports Medicine, 37(2) pp. 145-168.
Williams, A.G., Wilkinson, M. (2007) 'Simple Anthropometric and Physical Performance Tests to Predict Maximal Box-Lifting Ability.' The Journal of Strength and Conditioning Research, 21(2) pp. 638-638.
Little, T., Williams, A.G. (2007) 'Measures of Exercise Intensity During Soccer Training Drills With ProfessionalSoccer Players.' The Journal of Strength and Conditioning Research, 21(2) pp. 367-367.
Day, S.H., Gohlke, P., Dhamrait, S.S., Williams, A.G. (2007) 'No correlation between circulating ACE activity and VO
2max or mechanical efficiency in women.' European Journal of Applied Physiology, 99(1) pp. 11-18.Williams, A.G., Evans, P. (2007) 'Materials handling ability of regular and reserve British Army soldiers..' Mil Med, 172(2) pp. 220-223.
Berger, N.J.A., Rittweger, J., Kwiet, A., Michaelis, I., Williams, A.G., Tolfrey, K., Jones, A.M. (2006) 'Pulmonary O2 uptake on-kinetics in endurance- and sprint-trained master athletes..' Int J Sports Med, 27(12) pp. 1005-1012.
Williams, A.G., Rayson, M.P. (2006) 'Can simple anthropometric and physical performance tests track training-induced changes in load-carriage ability?.' Mil Med, 171(8) pp. 742-748.
Williams, A.G., Rayson, M.P. (2006) 'Can simple anthropometric and physical performance tests track training-induced changes in maximal box-lifting ability?.' Ergonomics, 49(7) pp. 661-670.
Little, T., Williams, A.G. (2006) 'Suitability of soccer training drills for endurance training.' Journal of Strength and Conditioning Research, 20(2) pp. 316-319.
Berger, N.J.A., Tolfrey, K., Williams, A.G., Jones, A.M. (2006) 'Influence of continuous and interval training on oxygen uptake on-kinetics..' Med Sci Sports Exerc, 38(3) pp. 504-512.
Little, T., Williams, A.G. (2006) 'Effects of differential stretching protocols during warm-ups on high-speed motor capacities in professional soccer players.' Journal of Strength and Conditioning Research, 20(1) pp. 203-207.
Little, T., Williams, A.G. (2006) 'Suitability of Soccer Training Drills for Endurance Training.' The Journal of Strength and Conditioning Research, 20(2) pp. 316-316.
Little, T., Williams, A.G. (2006) 'Effects of Differential Stretching Protocols During Warm-Ups on High-Speed Motor Capacities in Professional Soccer Players.' The Journal of Strength and Conditioning Research, 20(1) pp. 203-203.
Williams, A.G. (2005) 'Effects of basic training in the British army on regular and reserve army personnel (vol 19, pg 254, 2005).' JOURNAL OF STRENGTH AND CONDITIONING RESEARCH, 19(3) pp. 484-484.
Williams, A.G., Day, S.H., Folland, J.P., Gohlke, P., Dhamrait, S., Montgomery, H.E. (2005) 'Circulating angiotensin converting enzyme activity is correlated with muscle strength..' Med Sci Sports Exerc, 37(6) pp. 944-948.
Williams, A.G. (2005) 'Effects of basic training in the British Army on regular and reserve army personnel..' J Strength Cond Res, 19(2) pp. 254-259.
Little, T., Williams, A.G. (2005) 'Specificity of acceleration, maximum speed, and agility in professional soccer players.' Journal of Strength and Conditioning Research, 19(1) pp. 76-78.
Little, T., Williams, A.G. (2005) 'Specificity of Acceleration, Maximum Speed, and Agility in Professional Soccer Players.' The Journal of Strength and Conditioning Research, 19(1) pp. 76-76.
Williams, A.G. (2005) 'Effects of Basic Training in the British Army on Regular and Reserve Army Personnel.' The Journal of Strength and Conditioning Research, 19(2) pp. 254-254.
Day, S.H., Williams, C., Folland, J.P., Gohlke, P., Williams, A.G. (2004) 'The acute effects of exercise and glucose ingestion on circulating angiotensin-converting enzyme in humans.' European Journal of Applied Physiology, 92(4-5) pp. 579-583.
Williams, A.G., Dhamrait, S.S., Wootton, P.T.E., Day, S.H., Hawe, E., Payne, J.R., Myerson, S.G., World, M., Budgett, R., Humphries, S.E., Montgomery, H.E. (2004) 'Bradykinin receptor gene variant and human physical performance.' Journal of Applied Physiology, 96(3) pp. 938-942.
Williams, A.G., Rayson, M.P., Jones, D.A. (2004) 'Training diagnosis for a load carriage task..' J Strength Cond Res, 18(1) pp. 30-34.
Williams, A.G., Rayson, M.P., Jones, D.A. (2004) 'Training Diagnosis for a Load Carriage Task.' The Journal of Strength and Conditioning Research, 18(1) pp. 30-30.
Woods, D.R., World, M., Rayson, M.P., Williams, A.G., Jubb, M., Jamshidi, Y., Hayward, M., Mary, D.A.S.G., Humphries, S.E., Montgomery, H.E. (2002) 'Endurance enhancement related to the human angiotensin I-converting enzyme I-D polymorphism is not due to differences in the cardiorespiratory response to training.' European Journal of Applied Physiology, 86pp. 240-244.
Williams, A. (2002) 'Fatigue is not a necessary stimulus for strength gains during resistance training: Commentary.' British Journal of Sports Medicine, 36(5)
Williams, A.G., Day, S.H., Dhamrait, S., Fuentes, R.M. (2002) 'ACE gene, physical activity, and physical fitness.' Journal of Applied Physiology, 93(4) pp. 1561-1562.
Williams, A.G., Rayson, M.P., Jones, D.A. (2002) 'Resistance training and the enhancement of the gains in material-handling ability and physical fitness of British Army recruits during basic training.' Ergonomics, 45(4) pp. 267-279.
Williams, A.G., Ismail, A.N., Sharma, A., Jones, D.A. (2002) 'Effects of resistance exercise volume and nutritional supplementation on anabolic and catabolic hormones.' European Journal of Applied Physiology, 86pp. 315-321.
Williams, A.G., Day, S.H., Dhamrait, S. (2002) 'ACE gene, physical activity, and physical fitness [1] (multiple letters).' Journal of Applied Physiology, 93(4) pp. 1561-1562.
Williams, A.G., van den Oord, M., Sharma, A., Jones, D.A. (2001) 'Is glucose/amino acid supplementation after exercise an aid to strength training?.' British Journal of Sports Medicine, 35(2) pp. 109-113.
Williams, A.G., Rayson, M.P., Jubb, M., World, M., Woods, D.R., Hayward, M., Martin, J., Humphries, S.E., Montgomery, H.E. (2000) 'The ACE gene and muscle performance.' Nature, 403pp. 614-614.
Williams, A.G., Rayson, M.P., Jones, D.A. (1999) 'Effects of basic training on material handling ability and physical fitness of British Army recruits.' Ergonomics, 42(8) pp. 1114-1124.
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Non-peer reviewed articles / reviews
Armstrong, N., Ladlow, P., Smith, J., Williams, A.G. (2023) Enabling occupational groups to maintain their leading edge.
Stebbings, G., Herbert, A., Heffernan, S., Pielke Jr., R., Tucker, R., Williams, A. (2021) The BASES Expert Statement on Eligibility for Sex Categories in Sport: Trans Athletes.
Stebbings, G., Herbert, A., Heffernan, S., Pielke Jr., R., Williams, A. (2021) The BASES Expert Statement on Eligibility for Sex Categories in Sport: DSD Athletes.
Williams, A. (2019) Caster Semenya: Researcher an expert witness in Olympic champion's legal battle against IAAF rules.
Williams, A. (2019) Caster Semenya: The three aspects to consider on testosterone rules that do not justify discriminating against her.
Williams, A.G., Day, S.H., Stebbings, G., Erskine, (2017) What does ‘elite’ mean in sport and why does it matter?.
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Conference papers
Stebbings, G., Herbert, A., Pielke Jr., R., Tucker, R., Williams, A., Heffernan, S. (2021) 'The BASES Expert Statement on Eligibility for Sex Categories in Sport: Trans Athletes.' In BASES Conference 2021. 39. 16/11/2021 - 18/11/2021. Taylor & Francis, pp. 3-5.
Stebbings, G., Herbert, A., Heffernan, S., Pielke Jr., R., Williams, A. (2021) 'D3.S2.1(1) The BASES Expert Statement on Eligibility for Sex Categories in Sport: DSD Athletes.' In BASES Conference 2021. 39. 16/11/2021 - 18/11/2021. pp. 7-8.
Antrobus, M.R., Brazier, J., Herbert, A.J., Stebbings, G.K., Day, S.H., Heffernan, S.M., Kilduff, L.P., Cullen, T., Raleigh, S.M., Erskine, R.M., Williams, A. (2020) 'Association between COMT polymorphism but not BDNF-AS or NOS3 and elite rugby athlete status.' In 25th ECSS Anniversary Congress. Virtual, 28/10/2020 - 30/10/2020.
Brazier, J., Antrobus, M.R., Herbert, A.J., Stebbings, G.K., Day, S.H., Heffernan, S.M., Kilduff, L.P., Cullen, T., Raleigh, S.M., Erskine, R.M., Williams, A. (2020) 'Tendon and ligament-related genes associate with elite status in rugby.' In 25th ECSS Anniversary Congress. Virtual, 28/10/2020 - 30/10/2020.
Heffernan, S., Kilduff, L., Erskine, R., Day, S., Stebbings, G., Cook, C., Grainger, A., Brazier, J., Antrobus, M., Williams, A.G. (2018) 'Apolipoprotein E ε4 allele is not associated with elite rugby status but is present in 30% of athletes.' In 23rd Annual Congress of the European College of Sport Science. Dublin, Ireland, 4/7/2018 - 7/7/2018.
Antrobus, M., Brazier, J., Herbert, A., Stebbings, G., Day, S., Heffernan, S., Erskine, R., Kilduff, L., Williams, A.G. (2018) 'Association between MAPT polymorphism but not APOE promotor and elite rugby athlete status.' In 23rd Annual Congress of the European College of Sport Science. Dublin, Ireland, 4/7/2018 - 7/7/2018.
Brazier, J., Antrobus, M., Herbert, A., Stebbings, G., Day, S., Heffernan, S., Erskine, R., Kilduff, L., Williams, A.G. (2018) 'Association of MMP3 but not TIMP2 gene variants with elite rugby player status and rugby code.' In 23rd Annual Congress of the European College of Sport Science. Dublin, Ireland, 4/7/2018 - 7/7/2018.
Herbert, A., Williams, A.G., Lockey, S., Erskine, R., Sale, C., Hennis, P., Antrobus, M., Brazier, J., Heffernan, S., Day, S., Stebbings, G. (2018) 'Bone mineral density and associated genetic variants in high-level Caucasian marathon runners.' In 23rd Annual Congress of the European College of Sport Science. Dublin, Ireland, 4/7/2018 - 7/7/2018.
Herbert, A.J., Williams, A.G., Lockey, S.J., Erskine, R.M., Hennis, P.J., Sale, C., Day, S.H., Stebbings, G.K. (2018) 'Associations of bone mineral density-related genes and marathon performance in elite European Caucasian marathon runners..' In International Sports Science and Sports Medicine Conference (ISSMC). Newcastle University, 5/9/2017 - 7/9/2017.
Stebbings, G., Herbert, A.J., Brazier, J., Lockey, S.J., Erskine, R.M., Day, S.H., Williams, A.G. (2017) 'No association between tendon-related genes and performance in elite European Caucasian marathon runners..' In International Sports Science and Sports Medicine Conference (ISSMC). Newcastle University, 5/9/2017 - 7/9/2017.
Brazier, J.J., Antrobus, M.R., Heffernan, S.M., Stebbings, G.K., Day, S.H., Hennis, P.J., Erskine, R.M., Kilduff, L.P., Williams, A.G. (2017) 'An investigation into genetic associations with musculoskeletal soft tissue injuries in elite rugby – preliminary findings.' In 8th Annual Strength and Conditioning Student Conference. London,
Herbert, A.J., Williams, A.G., Lockey, S.J., Erskine, R.M., Heffernan, S.M., Pedlar, C.R., Kipps, C., Day, S.H., Stebbings, G.K. (2016) 'ACTN3 R577x genotype is not associated with elite european caucasian marathon performance.' 50. 29/9/2016 - 2/10/2016. BMJ Publishing Group, pp. A53-A54.
Stebbings, , Williams, A.G., Herbert, , Lockey, S.J., Heffernan, , Erskine, , Morse, C.I., Day, S.H. (2016) 'Titin genotype is associated with skeletal muscle fascicle length in recreationally active men and running performance in habitually trained marathon runners.' 50. 29/9/2016 - 2/10/2016. BMJ Publishing Group, pp. A55-A56.
Stebbings, G.K., Williams, A.G., Morse, C.I., Day, S.H. (2015) 'Association of TTN genotype with skeletal muscle fascicle length in untrained Caucasian males.' In Physiology 2015. 34. Cardiff,
Lockey, S.J., Erskine, R.M., Pedlar, C.R., Kipps, C.W.G., Heffernan, S.M., Herbert, A.J., Stebbings, G.K., Williams, A.G., Day, S.H. (2015) 'The individual and combined association between multiple gene polymorphisms and elite endurance athlete status.' In 20th ECSS Annual Congress. Malmo, Sweden,
Lockey, S.J., Erskine, R.M., Herbert, A.J., Pedlar, C.R., Kipps, C.W.G., Williams, A.G., Day, S.H. (2014) 'ACTN3 R577X genotype is not associated with elite marathon runners.' In Physiological Society. London, UK,
Stebbings, G.K., Morse, C.I., Williams, A.G., Day, S.H. (2014) 'ASSOCIATION OF ACTN3, CNTF AND PTK2 WITH SKELETAL MUSCLE PHENOTYPES IN UNTRAINED MALES.' In 19th Annual Congress of the European College of Sport Science (2014). Amsterdam,
Heffernan, S., Kilduff, L.P., Erskine, R.M., Day, S.H., McMahon, G.E., Cook, C.J., Vance, B., Wang, G., Collins, M., Pitsiladis, Y.P., Williams, A.G. (2013) 'ACE and ACTN3 gene variants are associated with elite status and playing position in rugby union.' In BASES 2013 Annual Conference. Preston, UK: BASES, pp. 74-75.
Stebbings, G.K., Morse, C.I., Williams, A.G., Day, S.H. (2013) 'Variability and distribution of muscle strength and its determinants in humans.' In BASES 2013 Annual Conference. Preston, UK: BASES, pp. 141-142.
Boyd, C., Barnes, C., Marsh, L., Williams, A.G. (2013) 'Time-motion analysis of football for athletes with cerebral palsy: special reference to classification.' In UK High Performance Conference for Paralympic Sports Science and Sports Medicine. Burton-upon-Trent, UK,
Ahmetov, I.I., Druzhevskaya, A.M., Lyubaeva, E.V., Popov, D.V., Vinogradova, O.L., Williams, A.G. (2011) 'ACTN3 genotype is associated with preferred racing distance and muscle fibre composition in speed skaters.' In ECSS 2011. Liverpool, UK,
Day, S.H., Williams, A.G., Turner, M., McPhee, J.S. (2011) 'The solute carrier family 22 gene rs2457571 polymorphism does not influence oxygen uptake responses to a six-week aerobic training programme.' In ECSS 2011. Liverpool, UK,
Williams, A.G., Day, S.H., Johnson, D., McPhee, J.S. (2011) 'The supervillin polymorphism rs6481619 is not associated with maximal oxygen uptake or its training response.' In ECSS 2011. Liverpool, UK,
Hughes, D.C., Williams, A.G. (2010) 'Muscle strength & power phenotypes: Polygenic profile similarity limits skeletal muscle performance.' In First Wingate Conference of Exercise and Sport Sciences. Wingate Institute, Israel,
McPhee, J.S., Degens, H., Jones, D.A., Baar, K., Williams, A.G. (2009) 'Individual variability in skeletal muscle metabolic response to endurance training in humans.' 14. 2009 - The Physiological Society, pp. 31-32.
McPhee, J.S., Jones, D.A., Williams, A.G. (2008) 'HIF1A genetic polymorphism and endurance performance.' In BASES 2008. 26. London, UK, Taylor & Francis: SSH Journals, pp. S18-S19.
McPhee, J.S., Tomlinson, D., Williams, A.G., Jones, D.A. (2008) 'Prescribing exercise intensity based on VO2max fails to induce peripheral training adaptations in all people.' In ECSS 2008. Lisbon, Portugal,
Williams, A.G., Folland, J.P. (2007) 'An algorithm for genetic potential to express muscle strength.' In ECSS 2007. Jyvaskyla, Finland,
Williams, A.G., Folland, J.P. (2007) 'Similarity of polygenic profiles limits the potential for elite human physical performance.' In ECSS 2007. Jyvaskyla, Finland,
Berger, N.J.A., Tolfrey, K., Williams, A.G., Jones, A.M. (2005) 'Effects of endurance training on the oxygen uptake-work rate slope during ramp exercise in humans.' In BASES 2004. 23. Liverpool, UK, Taylor & Francis: SSH Journals, pp. 135-136.
Little, T., Williams, A.G. (2004) 'Effects of differential stretching protocols during warm-ups on high speed motor capacities in professional footballers.' In Fifth World Congress of Science and Football. 22. Lisbon, Portugal, Taylor & Francis: SSH Journals, pp. 589-590.
Little, T., Williams, A.G. (2004) 'Specificity of acceleration, maximum speed and agility in professional soccer players.' In Fifth World Congress of Science and Football. 22. Lisbon, Portugal, Taylor & Francis: SSH Journals, pp. 550-551.
Wilkinson, M., Williams, A.G. (2003) 'Validity of anthropometric and strength tests for the prediction of box-lifting performance.' In BASES 2002 Annual Conference. 21. Manchester, Taylor & Francis: SSH Journals, pp. 309-310.
Bloomer, A., van Block, P., Williams, A.G. (2003) 'Influence of anthropometric measurements and date-of-birth on selection processes in youth rugby.' In Fifth International Conference in Sport, Leisure and Ergonomics. Liverpool, UK,
Wilkinson, M., Williams, A.G. (2002) 'Validity of simple field-based strength and anthropometric tests for the prediction of performance in a box-lifting task.' In BASES 2001. 20. Newport, UK, Taylor & Francis: SSH Journals, pp. 43-43.
Williams, A.G., Ismail, A.N., Sharma, A., Jones, D.A. (2001) 'Effects of resistance exercise volume and nutritional supplementation on anabolic and catabolic hormones.' In BASES 2000. 19. Wolverhampton, UK, Taylor & Francis: SSH Journals, pp. 646-646.
Williams, A.G., Ismail, A.N., Sharma, A., Jones, D.A. (2000) 'Effects of Resistance Exercise Volume and Nutritional Supplementation on Anabolic and Catabolic Hormones.' In 5th ECSS Congress. Jyvaskyla, Finland,
Williams, A.G., van den Oord, M., Jones, D.A. (2000) 'Is insulin an aid to resistance training?.' In BASES 1999. 18. Leeds, UK, Taylor & Francis: SSH Journals, pp. 551-551.
Williams, A.G., Rayson, M.P., Jones, D.A. (1999) 'Is training diagnosis for a load carriage task possible from simple pre-training tests?.' In BASES 1998. 17. Worcester, UK, Taylor & Francis: SSH Journals, pp. 21-22.
Williams, A.G., Rayson, M.P., Jones, D.A. (1999) 'Resistance training in the military enhances gains in lifting and loaded marching performance.' In BASES 1998. 17. Worcester, UK, Taylor & Francis: SSH Journals, pp. 22-22.
Williams, A.G., Rayson, M.P., Jones, D.A. (1999) 'The sensitivity of isometric and dynamic tests of strength in monitoring training-induced changes in lifting performance.' In BASES 1998. 17. Worcester, UK, Taylor & Francis: SSH Journals, pp. 23-23.
Williams, A.G., Irving, C.M.A., Sykes, K. (1999) 'Validation of the Tanita bioelectrical impedance body composition analyser.' In BASES 1998. 17. Worcester, UK, Taylor & Francis: SSH Journals, pp. 16-17.
Rayson, M.P., Williams, A.G. (1998) 'Do simple gym tests reflect training-induced changes in a lift task?.' In ACSM 1998. 30. Florida, USA, American College of Sports Medicine (ACSM), pp. S316-S316.
Williams, A.G., Lamb, K.L. (1997) 'Validation of two indirect methods of assessing body composition against hydrodensitometry with the emphasis on statistical methodology.' In BASES 1996. 15. Lilleshall, UK, Taylor & Francis: SSH Journals, pp. 35-35.
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Presentations
Williams, A.G., Mitra, P. (2021) Sex and gender boundaries in sport and exercise: A discussion on the science behind the sex-testing policies and the lived experiences of athletes. [Presentation] Virtual,16/11/2021.
Williams, A. (2021) The genetics of sex testing in sport. [Presentation] Athens, Greece,26/9/2021.
Williams, A. (2021) Genetophysiological research required at the male-female eligibility boundary. [Presentation] Athens, Greece,24/9/2021.
Williams, A. (2021) Sex-based sport performance differences and shortcomings of biological definitions. [Presentation] Paris, France,20/9/2021.
Williams, A. (2021) Are genetic tests in sport useful for practitioners?. [Presentation] Virtual, organised by Alexandria University, Egypt,3/4/2021.
Williams, A. (2020) Ethics of genetic sex testing in sport and associated medical interventions. [Presentation] Wuhan, China (virtual),29/10/2020.
Williams, A.G. (2019) Gender and eligibility in sport: Is it ever justifiable for sporting rules to prevent an athlete from competing within their gender category based on their physical attributes?. [Presentation] London, UK,9/10/2019.
Williams, A.G. (2019) Genetic contributions to motor ability: Can research inform sports practice?. [Presentation] Krakow, Poland,19/9/2019.
Williams, A.G. (2018) Personalised nutrition: The role of genetic testing. [Presentation] Newcastle, UK,18/12/2018.
Williams, A.G. (2018) DNA testing will become as common as (and probably more common than) measuring body composition. [Presentation] Birmingham, UK,11/5/2018.
Williams, A.G. (2018) DNA testing for personalised nutrition – revolution or fabrication?. [Presentation] Birmingham, UK,11/5/2018.
Williams, A.G. (2017) Can genetics predict performance in sport?. [Presentation] Newcastle, UK,7/9/2017.
Williams, A.G. (2017) Is there a genetic predisposition to concussion?. [Presentation] Newcastle, UK,5/9/2017.
Williams, A.G. (2017) Genetics in sport and exercise. [Presentation] London, UK,8/2/2017.
Williams, A.G. (2016) Genetics in rugby - injury risk, physical performance and playing position. [Presentation] Wrexham, UK,30/11/2016.
Williams, A.G. (2016) Athletes: Born or made. [Presentation] Manchester, UK,11/11/2016.
Williams, A.G. (2016) Choosing phenotypes in the GENESIS and ATHLOME projects. [Presentation] Ljubljana, Slovenia,2/10/2016.
Williams, A.G. (2016) The untapped potential of genomics in sport and exercise. [Presentation] London, UK,26/9/2016.
Williams, A.G. (2016) Can a genetic test unlock your athletic potential?. [Presentation] Cardiff, UK,30/1/2016.
Williams, A.G. (2015) GENESIS (GENomics of Elite Status In Sport) - A UK athlete DNA biobank. [Presentation] Northampton, UK,14/10/2015.
Williams, A.G. (2015) GENESIS (GENesis of Elite Status In Sport). [Presentation] Santorini, Greece,15/5/2015.
Williams, A.G. (2014) GENESIS (GENesis of Elite Athlete Status In Sport). [Presentation] London, UK,8/9/2014.
Williams, A.G. (2014) Genetics and rugby – data from a multi-dimensional team sport. [Presentation] Sao Paulo, Brazil,9/6/2014.
Williams, A.G. (2014) The GENetics of Elite Status In Sport (GENESIS) project. [Presentation] Swansea, UK,25/4/2014.
Williams, A.G. (2013) Athletes: Born or made. [Presentation] London, UK,3/12/2013.
Williams, A.G. (2013) Genes, your exercise performance and sporting ability. [Presentation] Oswestry, UK,8/11/2013.
Williams, A.G. (2013) Genetics and Welsh rugby. [Presentation] Swansea, UK,15/8/2013.
Williams, A.G. (2012) Sport and exercise genomics research: Trying to make a practical impact in sport. [Presentation] Kiev, Ukraine,4/10/2012.
Williams, A.G. (2012) Genomics research in sport and exercise. [Presentation] Cambridge, UK,11/7/2012.
Williams, A.G. (2012) Genomics as a practical tool in sport: Have we reached the starting line?. [Presentation] Sao Paulo, Brazil,25/6/2012.
Williams, A.G. (2012) Being a physiological and genetic outlier will help win gold in London. [Presentation] Cambridge, UK,10/5/2012.
Williams, A.G. (2012) Genetics of sport performance: Data from classical and molecular genetics. [Presentation] Kazan, Russia,20/3/2012.
Williams, A.G. (2011) DNA testing in sport. [Presentation] Manchester, UK,25/10/2011.
Williams, A.G. (2011) Genetic medalling. [Presentation] London, UK,7/6/2011.
Williams, A.G. (2011) Genetics of sport performance. [Presentation] Kazan, Russia,24/3/2011.
Williams, A.G. (2010) Genetics and sport performance. [Presentation] Paris, France,10/12/2010.
Williams, A.G. (2010) Genetics and elite sport: To win gold in 2012, do you have to choose the right parents?. [Presentation] Oswestry, UK,31/5/2010.
Williams, A.G. (2010) Genetics in sport and exercise science. [Presentation] Oxford, UK,8/5/2010.
Williams, A.G. (2010) Born to run. [Presentation] London, UK,15/4/2010.
Williams, A.G. (2009) Genetic research and testing in sport and exercise science. [Presentation] Frankfurt, Germany,6/2/2009.
Williams, A.G. (2009) The biomedical revolution and sport. [Presentation] Cambridge, UK,2/2/2009.
Williams, A.G. (2008) Gene doping in sport - relevant information from sport and exercise genetics. [Presentation] Florence, Italy,1/7/2008.
Williams, A.G. (2008) British Association of Sport and Exercise Sciences (BASES) Position Stand on Genetic Testing in Sport. [Presentation] Maastricht, The Netherlands,23/2/2008.
Williams, A.G. (2007) Genetics in sport and exercise science: What we know. [Presentation] Oxford, UK,25/9/2007.
Williams, A.G. (2007) Genetic research in sport and exercise science: What we know. [Presentation] Bath, UK,1/9/2007.
Williams, A.G. (2006) Science of genetic tests of performance. [Presentation] London, UK,28/9/2006.
Williams, A.G. (2005) Inter-individual variability. [Presentation] Stoke-on-Trent, UK,8/11/2005.
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Other
Williams, A.G., Folland, J.P. (2008) Sport genetics: physical performance potential is dependent on genome.
Williams, A.G. (2007) How genes limit our physical performance.
Williams, A.G., Wilkinson, M. (2002) Tapering and muscle fiber types.
Press and media
I have commented frequently via a range of media, and remain available to comment, on sex and gender in sport, elite physical performance, genetic technology in sport and exercise, as well as elite rugby performance and injury. Feel free to contact me.