Dr Huw Lloyd
Dr Huw Lloyd
Senior Lecturer in Wildlife Ecology
My profile
Biography
I am Senior Lecturer at the Department of Natural Sciences, where I have been working since 2011. Prior to joining Manchester Metropolitan University, I worked for two years as a Conservation Training Officer for a global bird conservation NGO and as a Guest Lecturer at the School of Biology, Newcastle University. I conducted my PhD research on the ecology and conservation of high-Andean Polylepis bird communities in Peru, and have also conducted research on numerous tropical forest bird communities including in cloud forest and lowland tropical forests in Peru, and in fragmented rainforest habitats in Indonesia. I conduct research primarily through building regional, national and international research relationships, helping early-career scientists from developed and developing countries internationalize their science. Since 2009 I have been developing conservation science leadership capacity in over 12 different conservation organizations and academic institutions in Peru, Brazil, China, Ethiopia, Nepal, Thailand and Tonga, providing academic support to undergraduate and postgraduate students, some of whom have gone on to become lecturers, whilst others have gone on the successfully complete or enrol in PhD programs. In the UK I work closely with organisations such as Cheshire Wildlife Trusts and Forestry England.
At MMU, I have recently finished serving as an elected member (Faculty of Science and Engineering) of the Academic Board between 2019 and 2021. I was field course coordinator for the Department of Natural Sciences for six years, and now am Departmental Health and Safety Coordinator, responsible for mentoring and guiding staff, PDRAs and PhD candidates in the development and completion of all field-based research risk assessments. I was external examiner for undergraduate and post-graduate degree courses at the School of Biological Sciences, University of Chester for four years.
My research partnerships in China, specifically with Beijing Normal University developed over the last ten years, now plays a role in MMU’s international strategy, aimed at increasing staff and student mobility between MMU and Chinese institutions, resulting in a university-wide Memorandum of Understanding signed by both institutions, to help increase collaboration across a number of faculties.
Interests and expertise
My main interests are wildlife ecology/biology and behaviour, conservation biology and ornithology. My primary research areas focus is on avian ecology, biology, evolution and conservation but I also work with bats in the UK and primates in Peru. I have particular interests in developing and testing survey methods, particularly acoustic monitoring of bird and bat populations, but also am interested in studies on avian foraging behaviour, factors influencing the outcome of conservation translocations, and understanding how bird species are likely to be impacted by further habitat/climate change.
In China, I have worked with colleagues from numerous universities on how migratory waterbird populations respond to natural, restored and artificial wetland environmental gradients; identifying key prey resources for threatened migratory birds; understanding the impacts of tourism on threatened migratory bird populations in coastal wetlands; quantifying how climate change is predicted to impact populations of forest-dependent bird species; how anthropogenic disturbance can create an ecological trap for habitat-dependent tidal reed-bed specialists; using bioacoustics for monitoring threatened bird species and improving our knowledge on the function of primary song types in species such as Eurasian Cuckoo; and developing new theoretical models on understanding the evolution of avian life-history traits. I have been involved in threatened species-poverty alleviation research in Ethiopia as part of a multidisciplinary research collaboration funded by the Darwin Initiative.
I am an Associate Editor for the open access journal Avian Research and Editorial Board member for the open access journal Watershed Ecology and the Environment, and currently Guest Editor for a special edition of the journal Diversity, on Avian Ecology, Diversity and Population Monitoring
Teaching
I am currently unit coordinator for two undergraduate units (Conservation Science in Practice, Wildlife Ecology and Behaviour) and one post-graduate unit (Avian Biology and Conservation), and teach across a number of other undergraduate and post-graduate units. For Conservation Science in Practice, I work with Cheshire Wildlife Trust to deliver an evidence-based taught unit with a novel assessment for the students based on the new Environment Bill, whilst the work of the trust is a central focus of the units activities (including lectures from the Director of Conservation and day visits to trust reserves). During my time at MMU I have designed two new residential field courses (South Wales and Portugal) that enable students to develop skills in both traditional biological field techniques, and also more contemporary remote sensing skills such as camera trapping and bioacoustics surveys, and the use of specialist software (e.g. Kaleidoscope, Program DISTANCE). Research-informed teaching is central to my lectures and to the teaching teams I work with. Research output from my international collaborations (e.g. wetland migratory bird ecology and behaviour; impacts of climate change on bird species distributions; threatened species translocations and bird conservation-poverty alleviation research) is utilised for both undergraduate and post-graduate teaching.
Research outputs
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Books (authored/edited/special issues)
Walker, B., Cheshire, G., Lloyd, H. (2007) Peruvian Wildlife A Visitor's Guide to the Central Andes. Bradt Travel Guides.
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Journal articles
Wu, X.Y., Lloyd, H., Dong, L., Zhang, Y.Y., Lyu, N. (2024) 'Moving beyond defining animal personality traits: The importance of behavioural correlations and plasticity for conservation translocation success.' Global Ecology and Conservation, 49
Liu, P., Lou, Y., Lloyd, H., Sun, Y. (2023) 'Multiple field study approaches reveal the functions of female song and duets in a temperate songbird.' Animal Behaviour, 206pp. 1-11.
Lou, Y., Zhao, Q., Hu, Y., Chen, L., Liu, P., Fang, Y., Lloyd, H., Sun, Y. (2023) 'Personality-dependent nest site selection and nest success during incubation in wild chestnut thrushes.' iScience, 26(8) pp. 107419-107419.
Lloyd, H. (2023) 'Editorial: Avian ecology and diversity, population monitoring and conservation: introduction to the Special Issue.' Diversity, 15(4) pp. 520-520.
Li, D., Bai, Y., Lei, W., Que, P., Liu, Y., Pagani-Núñez, E., Lloyd, H., Zhang, Z. (2023) 'Mammalian predators and vegetated nesting habitat drive reduced protected area nesting success of Kentish plovers, Yellow Sea region, China.' Ecology and Evolution, 13(3) pp. e9884-e9884.
Delaunay, M.G., Brassey, C., Larsen, C., Lloyd, H., Grant, R.A. (2022) 'The evolutionary origin of avian facial bristles and the likely role of rictal bristles in feeding ecology.' Scientific Reports, 12
Liu, P., Lloyd, H., Lou, Y., Sun, Y. (2022) 'Soft song provokes stronger aggressive responses than broadcast song in the Plain Laughingthrush (Pterorhinus davidi).' IBIS: The International Journal of Avian Science, 165(2) pp. 561-570.
Lyu, N., Yeh, D.J., Lloyd, H., Sun, Y.H. (2022) 'The significant role of post-pairing male behavior on the evolution of male preferences and female traits.' Communications Biology, 5
Lyu, N., Hu, Y., Zhang, J., Lloyd, H., Sun, Y.H., Tao, Y. (2021) 'Switching costs in stochastic environments drive the emergence of matching behaviour in animal decision-making through the promotion of reward learning strategies.' Scientific Reports, 11(1)
Yang, C., Wang, J., Lyu, N., Lloyd, H. (2021) 'Comparison of digital photography and spectrometry for evaluating colour perception in humans and other trichromatic species.' Behavioral Ecology and Sociobiology, 75(11)
Li, D., Han, M., Lloyd, H., Jin, L., Zhang, L., Yin, J., Wan, D. (2021) 'Loss of extra-pair paternity is not associated with decreased paternal investment in mixed-paternity broods or unrelated nestlings in the varied tit, Parus varius.' Pakistan Journal of Zoology, 53(6) pp. 2105-2116.
Zhang, J., Zhang, Y., Lloyd, H., Zhang, Z., Li, D. (2021) 'Rapid reclamation and degradation of suaeda salsa saltmarsh along coastal china’s northern yellow sea.' Land, 10(8) pp. 835-835.
Zhang, J., Zhang, H., Liu, Y., Lloyd, H., Li, J., Zhang, Z., Li, D. (2021) 'Saltmarsh vegetation and social environment influence flexible seasonal vigilance strategies for two sympatric migratory curlew species in adjacent coastal habitats.' Avian Research, 12(1)
Bedson, C.P.E., Thomas, L., Wheeler, P.M., Reid, N., Harris, W.E., Lloyd, H., Mallon, D., Preziosi, R. (2021) 'Estimating density of mountain hares using distance sampling: a comparison of daylight visual surveys, night-time thermal imaging and camera traps.' Wildlife Biology, 2021(3)
Mahamued, B.A., Donald, P.F., Collar, N.J., Marsden, S.J., Ndang'ang'a, P.K., Wondafrash, M., Abebe, Y.D., Bennett, J.A.M.E.S., Wotton, S.R., Gornall, D., Lloyd, H.U.W. (2021) 'Rangeland loss and population decline of the critically endangered Liben Lark Heteromirafra archeri in southern Ethiopia.' Bird Conservation International, 32(1) pp. 64-77.
Wu, F., Lei, W., Lloyd, H., Zhang, Z. (2020) 'Predictors of Gull-billed tern (Gelochelidon nilotica) nest survival in artificial coastal saltpans, Bohai Bay, China.' PeerJ, 8pp. e10054-e10054.
Li, D., Zhang, J., Chen, L., Lloyd, H., Zhang, Z. (2020) 'Burrow ambient temperature influences Helice crab activity and availability for migratory Red-crowned cranes Grus japonensis.' Ecology and Evolution, 10(20) pp. 11523-11534.
Li, D., Zhang, J., Liu, Y., Lloyd, H., Pagani-Núñez, E., Zhang, Z. (2020) 'Differences in dietary specialization, habitat use and susceptibility to human disturbance influence feeding rates and resource partitioning between two migratory Numenius curlew species.' Estuarine, Coastal and Shelf Science, 245pp. 106990-106990.
Delaunay, M.G., Larsen, C., Lloyd, H., Sullivan, M., Grant, R.A. (2020) 'Anatomy of avian rictal bristles in Caprimulgiformes reveals reduced tactile function in open-habitat, partially diurnal foraging species.' Journal of Anatomy, 237(2) pp. 355-366.
Xia, C., Deng, Z., Lloyd, H., Moller, A.P., Zhao, X., Zhang, Y. (2019) 'The function of three main call types in common cuckoo.' Ethology, 125(9) pp. 652-659.
Deng, Z., Lloyd, H., Xia, C., Li, D., Zhang, Y. (2019) 'Within-season decline in the call consistency of individual male Common Cuckoo (Cuculus canorus).' Journal of Ornithology, 160(2) pp. 317-327.
Deng, Z., Lloyd, H., Xia, C., Møller, A., Liang, W., Zhang, Y. (2018) 'Components of variation in female common cuckoo calls.' Behavioural Processes, 158pp. 106-112.
Mendez, D., Marsden, S., Lloyd, H. (2018) 'Assessing population size and structure for Andean Condor Vultur gryphus in Bolivia using a photographic ‘capture‐recapture’ method.' Ibis, 161(4) pp. 867-877.
Xia, C., Lloyd, H., Shi, J., Wei, C., Zhang, Y. (2018) 'Dawn singing of the Brownish‐flanked Bush Warbler influences dawn chorusing in a bird community.' Ethology, 124(6) pp. 400-409.
Li, D., Liu, Y., Sun, X., Lloyd, H., Zhu, S., Zhang, S., Wan, D., Zhang, Z. (2017) 'Habitat-dependent changes in vigilance behaviour of Red-crowned Crane influenced by wildlife tourism.' Scientific Reports, 7
Li, Y., Xia, C., Lloyd, H., Li, D., Zhang, Y. (2017) 'Identification of vocal individuality in male cuckoos using different analytical techniques.' Avian Research, 8(1)
Lyu, N., Servedio, M.R., Lloyd, H., Sun, Y.H. (2017) 'The evolution of post-pairing male mate choice.' Evolution, 71(6) pp. 1465-1477.
Boye, L., Li, L., Lloyd, H., Canwei, X., Zhang, Y., Zheng, G. (2016) 'Comparing post-release survival and habitat use by captive-bred Cabot’s Tragopan (Tragopan caboti) in an experimental test of soft-release reintroduction strategies.' Avian Research, 7
Nan, Y., Moermond, T., Lloyd, H., Yu, X., Liang, D., Kai, Z., Bisong, Y., Jianghong, R. (2016) 'Effects of Supplementary Feeding on the Breeding Ecology of the Buff-Throated Partridge in a Tibetan Sacred Site, China.' PLoS One, 11(1) pp. e0146568-e0146568.
Lyu, N., Lloyd, H., Sun, Y.H. (2015) 'Delayed plumage maturation in birds and the significance of condition-dependent parental care.' Behavioral Ecology and Sociobiology,
Lyu, N., Lloyd, H., Sun, Y.H. (2015) 'Delayed plumage maturation in birds and the significance of condition-dependent parental care.' Behavioral Ecology and Sociobiology, 69(6) pp. 1003-1010.
Xia, C., Liu, B., Wang, D., Lloyd, H., Zhang, Y. (2015) 'Reliability of the Brownish-flanked Bush Warbler’s soft song in male-male conflict.' Avian Research, 6
Li, D., Sun, X., Lloyd, H., Que, P., Liu, Y., Wan, D., Zhang, Z. (2015) 'Reed Parrotbill nest predation by tidal mudflat crabs: Evidence for an ecological trap?.' ECOSPHERE, 6
Li, D., Ding, Y., Yuan, Y., Lloyd, H., Zhang, Z. (2014) 'Female tidal mudflat crabs represent a critical food resource for migratory Red-crowned Cranes in the Yellow River Delta, China.' Bird Conservation International, 24(4) pp. 416-428.
LI, D., DING, Y., YUAN, Y., LLOYD, H., ZHANG, Z. (2014) 'Female tidal mudflat crabs represent a critical food resource for migratory Red-crowned Cranes in the Yellow River Delta, China.' Bird Conservation International,
Xia, C., Wei, C., Lloyd, H., Liu, J., Wu, Q., Zhang, Y. (2014) 'Dawn singing intensity of the male brownish-flanked bush warbler: Effects of territorial insertions and number of neighbors.' Ethology, 120(4) pp. 324-330.
Torres-Sovero, C., Lloyd, H., Brightsmith, D.J. (2014) 'Clay lick use and other observations of the Blue-headed Macaw (Primolius couloni) in southeastern Peru.' Ornitologia Neotropical, 25(1) pp. 87-94.
Li, D., Chen, S., Lloyd, H., Zhu, S., Shan, K., Zhang, Z. (2013) 'The importance of artificial habitats to migratory waterbirds within a natural/artificial wetland mosaic, Yellow River Delta, China.' Bird Conservation International, 23(2) pp. 184-198.
Xia, C., Lin, X., Liu, W., Lloyd, H., Zhang, Y. (2012) 'Acoustic identification of individuals within large avian populations: a case study of the Brownish-flanked Bush Warbler, South-Central China..' PLoS One, 7(8)
Lu, N., Jia, C.X., Lloyd, H., Sun, Y.H. (2012) 'Species-specific habitat fragmentation assessment, considering the ecological niche requirements and dispersal capability.' Biological Conservation, 152pp. 102-109.
Zhou, Z., Sun, Y., Dong, L., Xia, C., Lloyd, H., Zhang, Y. (2012) 'Breeding biology of Asian House Martin Delichon dasypus in a high-elevation area.' Forktail, (28) pp. 62-66.
Lloyd, H., Sevillano Ríos, S., Marsden, S.J., Valdés-Velásquez, A. (2012) 'Bird community composition across an Andean tree-line ecotone.' Austral Ecology, 37(4) pp. 470-478.
Lu, N., Jing, Y., Lloyd, H., Sun, Y.H. (2012) 'Assessing the distributions and potential risks from climate change for the Sichuan Jay (perisoreus internigrans).' Condor, 114(2) pp. 365-376.
Bernardo, C.S.S., Lloyd, H., Olmos, F., Cancian, L.F., Galetti, M. (2011) 'Using post-release monitoring data to optimize avian reintroduction programs: A 2-year case study from the Brazilian Atlantic Rainforest.' Animal Conservation, 14(6) pp. 676-686.
Bernardo, C.S.S., Cresswell, B., Lloyd, H., Azeredo, R., Simpson, J. (2011) 'Selection of radio transmitter and attachment method for post-release monitoring of captive-bred reintroduced red-billed curassow Crax blumenbachii, Brazil.' European Journal of Wildlife Research, 57(3) pp. 689-694.
Ríos, S.S., Lloyd, H., Valdés-Velásquez, A. (2011) 'Bird species richness, diversity and abundance in Polylepis woodlands, Huascaran biosphere reserve, Peru.' Studies on Neotropical Fauna and Environment, 46(1) pp. 69-76.
Zhang, K., Yang, N., Xu, Y., Ran, J., Lloyd, H., Yue, B. (2011) 'Nesting behavior of Szechenyi's monal-partridge in treeline habitats, Pamuling Mountains, China.' Wilson Journal of Ornithology, 123(1) pp. 93-96.
Li, D., Chen, S., Guan, L., Lloyd, H., Liu, Y., Lv, J., Zhang, Z. (2011) 'Patterns of waterbird community composition across a natural and restored wetland landscape mosaic, Yellow River Delta, China.' Estuarine, Coastal and Shelf Science, 91(2) pp. 325-332.
Lloyd, H., Marsden, S.J. (2011) 'Between-Patch Bird Movements within a High-Andean Polylepis Woodland/Matrix Landscape: Implications for Habitat Restoration.' Restoration Ecology, 19(1) pp. 74-82.
Wei, M., Lloyd, H., Zhang, Y. (2011) 'Neighbour-stranger discrimination by Yellow-bellied Tit Parus venustulus: Evidence for the "dear-enemy" effect.' Journal of Ornithology, 152(2) pp. 431-438.
Lloyd, H., Bernardo, C.S.S., Bayly, N., Galetti, M. (2011) 'Modelling post-release survival of reintroduced Red-billed Curassows Crax blumenbachii, Brazil.' Ibis, 153pp. 562-572.
Lloyd, H. (2008) 'Influence of within-patch habitat quality on high-Andean Polylepis bird abundance.' Ibis, 150(4) pp. 735-745.
Lloyd, H., Marsden, S.J. (2008) 'Bird community variation across Polylepis woodland fragments and matrix habitats: Implications for biodiversity conservation within a high Andean landscape.' Biodiversity and Conservation, 17(11) pp. 2645-2660.
Lloyd, H. (2008) 'Foraging ecology of High Andean insectivorous birds in remnant Polylepis forest patches.' Wilson Journal of Ornithology, 120(3) pp. 531-544.
Lloyd, H. (2008) 'Abundance and patterns of rarity of rarity of Polylepis birds in the Cordillera Vilcanota, southern Perú: Implications for habitat management strategies.' Bird Conservation International, 18(2) pp. 164-180.
Lloyd, H. (2004) 'Habitat and population estimates of some threatened lowland forest bird species in Tambopata, south-east Peru.' Bird Conservation International, 14(4) pp. 261-277.
Lloyd, H. (2003) 'Population densities of some nocturnal raptor species (Strigidae) in southeastern Peru.' Journal of Field Ornithology, 74(4) pp. 376-380.
Lloyd, H. (2000) 'Population densities of the Black-faced Cotinga Conioptilon mcilhennyi in south-east Peru.' Bird Conservation International, 10(3) pp. 277-285.